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Yellow Tangle-vein Flies - Trichophthalma sp.

Family Nemestrinidae 

This page contains pictures and information about Yellow Tangle-vein Flies that we found in the Brisbane area, Queensland, Australia
Body length 15mm
The fly is golden yellowish-brown in colour, with large pointy abdomen. Its body is covered with short hairs. Its large pair of dark red compound eyes covered most of its head. It has the long mouth parts. Its pair of wings is tinted in brown.   
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At first sight in the field we thought this was a Beefly. Went back home and looked at the photos, from its complex wing veins, we recognized it was a Tangle-vein Fly.   
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We found this fly once in Daisy Hill near Buhot Creek during mid summer Dec 2008. 
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This Tangle-vein Fly has seven nearly parallel branches of veins reaching the anterior margin of the wing, 1 subcostal, 4 radials, and 2 medians. Veins rl and r5 are bifurcated. The fly has the long proboscis same length as its head. The dark red eyes are covered with dense and long hairs. Those confirmed the fly is Trichophthalma sp..
On wing the veinlet m3+4 present which makes the continuation of diagonal vein.
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This fly did not mind we came very close. It were hovering most of the time above short plants. Sometimes even hovering still in the air for s few seconds. Its landed on plants for one or two seconds then hovering again.
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On Feb 2013 in Karawatha Forest, we found one Yellow Tangle-vein Fly was captured by a Fiery Skimmer Dragonfly

1. Insects of Australia - CSIRO, Division of Entomology, Melbourne University Press, 2nd Edition 1991, p755.
2. Insects of Australia and New Zealand - R. J. Tillyard, Angus & Robertson, Ltd, Sydney, 1926, p360.
3. A review of Australian Nemestrinidae (Diptera) -SJ Paramonov, Australian Journal of Zoology 1(2) 242 - 290, CSIRO 1953. 

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Last updated: March 02, 2013.